School of Natural Sciences and Engineering, Reproductive biology and morphology of plants,Full Professor

of School Natural Sciences and Engineering,of Plant Genetic Resources, Institute of Botany, ISU

Maia Akhalkatsi graduated from Faculty of Biology of the Tbilisi State University in 1981.PhD thesis in botany: “The ultrastructure of trichomes of Urticaceae family members” defended from Komarov Botanical Institute (St. Petersburg, Russia) in 1986. Worked as a researcher at the National Academy of Sciences of Georgia in the Institute of Botany since 1985 and she is head of department of Plant Genetic Resources since 2002. Chairman of NGO Georgian Society of Nature Explorers Orchis” (GSNE) in 1997-2011. In 1997, graduated the certificate course as postdoctoral researcher in International Research and Exchanges Board (IREX), Portland University, Portland, Oregon, USA. Alexander von Humboldt Foundation Fellowship in 1997-1999, Heinrich Heine University, Düsseldorf, Germany. Since 1997 is a member of the Botanical Society of the United States and Germany. In 2006 defended Doctor of Biological Science thesis “Structural and functional aspects of plant reproductive strategies” at the Institute of Zoology, National Academy of Sciences of Georgia. The major lines of research: plant anatomy, morphology and reproduction, biodiversity, plant genetic resources, etc. Published 88 research papers. In Ilia State University works since 2006. Since 2015 is a council member of School Natural Sciences and Engineering and founder of Botanical Society of Georgia.

Scientific interests / research interests

  • Plant anatomy
  • Plant morphology
  • Plant reproductive biology
  • Biodiversity
  • Plant conservation
  • Plant genetic resources
  • Crop wild relatives
  • Domestication of cultivars
  • Medicinal plants
  • Microscopy

Featured publications (23)

  • Goloshvili Tamar, Badridze Gulnara, Akhalkatsi Maia. Characterization of grape seed extracts of native to Georgia varieties of Vitis vinifera L. Pakistan Journal of Botany 50 (1): 245-250. (Impact Factor: 0.947).
  • Winkler M., Lamprecht A., Steinbauer K., Hülber K., Theurillat J.-P., Breiner F., Choler Ph., Ertl S., Gutiérrez Girón A., Rossi G., Vittoz P., Maia Akhalkatsi, Bay Chr., Benito J. L., Bergström T., Carranza M. L., Corcket E., Dick J., Erschbamer B., Fernández Calzado R., Fosaa A. M., Gavilán R., Ghosn D., Gigauri Khatuna, Huber D., Kanka R., Kazakis G., Klipp, Jozef Kollar, Thomas Kudernatsch, Per Larsson, Martin Mallaun, Ottar Michelsen, Pavel Moiseev M., Moiseev D., Molau U., Mesa J. M.. di Cella U. M., Nagy L., Petey M., Puscas M., Rixen Ch., Stanisci A., Suen M., Syverhuset A. O., Tomaselli M., Unterluggauer P., Ursu T., Villar L., Gottfried M., Pauli H. 2016. The rich sides of mountain summits – a pan-European view on aspect preferences of alpine plants. Journal of Biogeography 43 (11): 2261–2273. DOI: 10.1111/jbi.12835. (Impact Factor: 4.248).
  • Gogniashvili Mari, Jinjikhadze Tamar, Maisaia Inesa, Akhalkatsi Maia, Kotorashvili Adam, Kotaria Nato, Beridze Tengiz, Dudnikov Alexander Ju.  Complete chloroplast genomes of Aegilops tauschiiCoss. and Ae. cylindrica Host sheds light on plasmon D evolution. Current Genetics 62 (4): 791–798. DOI: 10.1007/s00294-016-0583-5. (Impact Factor: 2.76).
  • Gigauri Kh., Akhalkatsi M., Nakhutsrishvili G., Abdaladze O. Alpine Plant Distribution and Thermic Vegetation Indicator on Gloria Summits in the Central Greater Caucasus. Pakistan Journal of Botany 48 (5): 1893-1902. (Impact Factor: 0.947).
  • Togonidze N., Akhalkatsi M. Variability of plant species diversity during the natural restoration of the subalpine birch forest in the Central Great Caucasus. Turkish Journal of Botany 39 (3): 458-471; DOI:10.3906/bot-1404-19. (Impact Factor: 1.178).
    • Asanidze Z., Akhalkatsi M., Henk A. D., Richards Ch. M., Volk G. M. Genetic relationships between wild progenitor pear (Pyrus L.) species and local cultivars native to Georgia, South Caucasus. Flora 209 (9): 504-512. DOI: 10.1016/j.flora.2014.06.013. (Impact Factor: 1.125).
    • Ekhvaia J., Gurushidze M., Blattner F.R., Akhalkatsi M. Genetic diversity of Vitis vinifera in Georgia: relationships between local cultivars and wild grapevine, V. vinifera subsp. sylvestrisGenetic Resources and Crop Evolution 61 (8): 1507-1521. DOI: 10.1007/s10722-014-0125-2. (Impact Factor: 1.43).
    • Abramidze T.,   Bragvadze T., Chikhelidze N., Akhalkatsi M., Gotua M., Gamkrelidze A. 2014 SEP. Seasonal distribution of airborne pollen and spores in Tbilisi, Georgia. ALLERGY  69 Supplement: 99: 329-330 Meeting Abstract: 867. (Impact Factor:995).
    • Gigauri Kh., Akhalkatsi M., Nakhutsrishvili G., Abdaladze O. Monitoring of vascular plant diversity in a changing climate in the alpine zone of the Central Greater Caucasus. Turkish Journal of Botany 37 (6): 1104-1114. DOI:10.3906/bot-1301-38.(Impact Factor: 1.178).

Pauli H., Gottfried M., Dullinger S., Abdaladze O., Akhalkatsi M., Alonso J.L.B., Coldea, Dick J., Erschbamer B., Fernández Calzado R., Ghosn D., Holten J.I., Kanka R., Kazakis G., Kollár J., Larsson P., MoiseevP., MoiseevD., Molau U., Molero Mesa J., Nagy L., Pelino G., Puşcaş M., Rossi G., Stanisci A., Syverhuset A.O., Theurillat J.P., Tomaselli M., Unterluggauer P., Villar L., Vittoz P., Grabherr G. 2012. Recent plant diversity trends on Europe’s mountain summits. Science 336: 353-355. DOI: 10.1126/science.1219033. (Impact Factor: 37.205).

  • Gottfried M., Pauli H., Futschik A., Akhalkatsi M., Barančok P., Alonso J.L.B., Coldea G., Dick J., Erschbamer B., Rosa M., Calzado F., Kazakis G., Krajči J., Larsson P., Mallaun M., Michelsen O., Moiseev D., Moiseev P., Molau U., Merzouki  , Nagy L., Nakhutsrishvili G., Pedersen B., Pelino G., Puscas M., Rossi G., Stanisci A., Theurillat J.-P., Tomaselli M., Villar L., Vittoz P., Vogiatzakis I., Grabherr G. 2012. An early warning signal for a continental-scale response to ongoing climate change. Nature Climate Change 2: 111–115. DOI: 10.1038/NCLIMATE1329. (Impact Factor: 19.304).
  • Asanidze Z., Akhalkatsi M., Gvritishvili M. Comparative morphometric study and relationships between the Caucasian species of wild pear (Pyrus spp.) and local cultivars in Georgia. Flora 206: 974-986. DOI:10.1016/j.flora.2011.04.010. (Impact Factor: 1.125).
  • Akhalkatsi M., Ekhvaia J., Mosulishvili M., Nakhutsrishvili G., Abdaladze O., Batsatsashvili K. Reasons and processes leading to the erosion of crop genetic diversity in mountainous regions of Georgia. Mountain Research and Development 30 (3): 304-310. DOI:10.1659/MRD-JOURNAL-D-10-00022.1. (Impact Factor: 1.03).
  • Ekhvaia J., Akhalkatsi M. 2010. Morphological variation and relationships of Georgian populations of Vitis vinifera subsp. sylvestris (C.C. Gmel.) Hegi. Flora 205: 608-617. doi:10.1016/j.flora.2009.08.002. (Impact Factor: 1.125).
  • Erschbammer B., Mallaun M., Unterluggauer P., Abdaladze O., Akhalkatsi M., Nakhutsrishvili G. Species richness and distribution above the treeline in the Central Alps (Italy) and in the Central Caucasus (Georgia). Tuexenia 30:11-29. (Impact Factor: 1.04).
  • Pistrick K., Akhalkatsi M., Girgvliani T., Shanshiashvili T. 2009. Collecting plant genetic resources in Upper Svaneti (Georgia, Caucasus Mountains).  Journal of agriculture and rural development in the tropics and subtropics, Supplement 92: 127-135. (Impact Factor: 67).
  • Hughes N.M., Johnson D.M., Akhalkatsi M., Abdaladze O. 2009. Characterizing Betula litwinowii seedling microsites at the alpine-treeline ecotone, Central Greater Caucasus Mountains, Georgia. Arctic, Antarctic, and Alpine Research 41: 112-118. DOI:10.1657/1938-4246(08-021)[HUGHES]2.0.CO;2. (Impact Factor: 74).
  • Akhalkatsi M., Abdaladze O., Nakhutsrishvili G., Smith W.K. Rhododendron caucasicum and microtopography extend the Betula litwinowii alpine treeline (Caucasus Mountains, Georgia)Arctic, Antarctic, and Alpine Research 38: 481-488. DOI:10.1657/1523-0430(2006)38[481:FOSMBR]2.0.CO;2. (Impact Factor: 1.74).
  • Akhalkatsi M., Lösch RWater limitation effect on seed development and germination in Trigonella coerulea (Fabaceae). Flora 200 (6): 493-501. doi:10.1016/j.flora.2005.06.001. (Impact Factor: 1.125).
  • Akhalkatsi M., Lösch R. Changes in Water Relations, Solute Leakage and Growth Characters During Seed Germination and Seedling Development in Trigonella coerulea (Fabaceae)Journal of Applied Botany and Food Quality 75 (3-4): 144-151. (Impact Factor: 1.20).
  • Akhalkatsi M., Pfauth M., Calvin C. Structural aspects of seed and ovule development and non-random abortion in Melilotus officinalis (Fabaceae). Protoplasma 208 (1-4): 211-223. DOI:10.1007/BF01279092. (Impact Factor: 1.68).
  • Akhalkatsi M., Wagner J. Comparative embryology of three mountain Gentianaceae in the Central Caucasus and the European Alps.Plant Systematics and Evolution 204 (3-4): 39-48. DOI:10.1007/BF00982530. (Impact Factor: 1.154).
  • Akhalkatsi M., Wagner J. Reproductive phenology and seed development of Gentianella caucasea in different habitats in the Central Caucasus. Flora 191: 161-168. (Impact Factor: 1.125).

Books (Georgian-1; Georgian-English-2)

  • Akhalkatsi Maia. 2017. Structural and functional aspects of plant reproductive strategy. Tbilisi. (Georg.)
  • Akhalkatsi M. 2015. Forest Habitat Restoration in Georgia, Caucasus Ecoregion. Mtsignobari, Tbilisi. pp.115. (Georg.-Eng.)
  • Akhalkatsi M., Kimeridze M., Künkele S., Lorenz R., Mosulishvili M. 2003. Diversity and conservation of Georgian orchids. Tbilisi, CGS Ltd. pp. 40. (Georg.-Eng.)
  • Book chapters (Georgian-English-1; English-5)
  • Akhalkatsi M. 2015. Crop wild related species in Georgia. In: Agrobiodiversity of georgia (Catalog); Autor and editor: Aleksidze, Georgia Academy of Agricultural sciences pp. 183-216. ISBN 978-9941-0-8245-0. Tbilisi. (Georg.-Eng.).
  • Akhalkatsi M. Erosion and Prevention of Crop Genetic Diversity Landraces of Georgia (South Caucasus). Genetic Diversity and Erosion in Plants, Springer International Publishing; 12/2015; pp.159-187.
  • Akhalkatsi 2014.Traditional viticulture and grapevine diversity in Georgia. In: Parolly G., Grotz K., Walter H. (eds). Caucasus. Plant Diversity between the Black and Caspian Seas. ISBN: 978-3-921800-90-4. BGBM Press, Berlin-Dahlem.
  • Akhalkatsi M., Ekhvaia J., Asanidze Z. Diversity and Genetic Erosion of Ancient Crops and Wild Relatives of Agricultural Cultivars for Food: Implications for Nature Conservation in Georgia (Caucasus). In: John Tiefenbacher (Ed.), Perspectives on Nature Conservation, ISBN: 978-953-51-0033-1, InTech, Croatia. pp. 51-92.
  • Nakhutsrishvili G., Abdaladze O., Akhalkatsi M. Biotope types of the treeline of the Central Greater Caucasus. In: D. Gafta, J. Akeroyd (eds), Nature Conservation: Concepts and Practice. Springer, Berlin, NY, pp. 211-225.
  • Akhalkatsi M. 2006. International cooperative projects in botany with participation of Georgian scientists. Georgien-Postsowjetische Umbrüche und Hoffnungen auf Europa. Alexander von Humboldt Stiftung – Arbeits- und Diskussiospapier 6: 37-39.
  • Monographs (5)
  • Akhalkatsi, M., Tarkhnishvili, D. Habitats of Georgia, GTZ, Tbilisi. (Georg.-Eng.)
  • Akhalkatsi, M. Habitats of Georgia, GTZ, Tbilisi. (Georg.-Eng.)
  • Akhalkatsi M. 2009. Conservation and sustainable use of crop wild relatives in Samtskhe-Javakheti. Tbilisi. Elkana.
  • Akhalkatsi Maia; Kimeridze Mariam; Mosulishvili Marine; Maisaia Inesa Conservation and Sustainable Utilization of Rare Medicinal Plants in Samtskhe-Javakheti. Final Report. Tbilisi pp. 167. (Georg.-Eng.)
  • Akhalkatsi, Kimeridze, M., Mosulishvili, M., Maisaia, I. 2005. Conservation and Sustainable Utilization of the Endangered Medicinal Plants in Samtskhe-Javakheti. Environmental Baseline. Tbilisi. pp. 80. (Georg.-Eng.)

Articles in magazines

  • Akhalkatsi Maia, Girgvliani Tamar. Crop Wild Relatives of Genus Coriandrum L. in Georgia (South Caucasus). Agricultural Research & Technology: Open Access Journal (ARTOAJ) 7 (3): 1-5. DOI:10.19080/ARTOAJ.2017.07.555712.
  • Akhalkatsi Maia; Otte Annette; Togonidze Natalia; Bragvadze Tamar; Asanidze Zezva; Arabuli Giorgi; Chikhelidze Natia; Mazanishvili Lamar. Agrobiodiversity and genetic erosion of crop varieties and plant resources in the Central Great Caucasus. Annals of Agrarian Science 15 (1)11–16.
  • Akhalkatsi Maia. 2017. Climate Global Change on Reproduction and Diversity of Agricultural Plants in Semi-Arid Regions of Georgia (Caucasus Ecoregion). Agricultural Research & Technology: Open Access Journal (ARTOAJ) 3 (4): 1-2DOI:10.19080/ARTOAJ.2017.03.5556
  • Akhalkatsi Maia. Ancient Crops Continuing for an Extended Period in Samtskhe-Javakheti Region of Georgia – A Review. Agricultural Research & Technology: Open Access Journal (ARTOAJ) 2 (5): 1-9DOI10.19080/ARTOAJ.2016.03.555605.
  • Akhalkatsi M, Gvaladze G, Girgvliani T. 2016. Situational Analyses of the Current State of Plant Genetic Resources Important for Food and Agriculture in Georgia (Caucasus Ecoregion). Agricultural Research & Technology: Open Access Journal (ARTOAJ) 2 (2): 1-5. DOI: 10.19080/ARTOAJ.2016.01.555582.
  • Akhalkatsi M, Girgvliani T. Landraces and Wild Species of the Secale Genus in the Georgia (Caucasus Ecoregion). Agricultural Research & Technology: Open Access Journal (ARTOAJ) 1 (4): 1-7. DOI10.19080/ARTOAJ.2016.01.555567.
  • Richard Lorenz, Maia Akhalkatsi, Pierluigi Cortis, Rosario Galesi, Cesario Giotta, Johann Madl, Erich Obrist, Marcello Piccitto, Vito Antonio Romano, Rolando Romolini und Romieg Soca. Morphometrische Untersuchungen zur Variabilitat und Gliederung der Gattung Platanthera in Italien. Journal Europäischer Orchideen 47 (1): 123-238.
  • Akhalkatsi M., Asanidze Z. Pyrus caucasica Fed. Ancestor of Georgian Pear Landraces. Science, Technology and Development 35 (2): 105-108. DOI:10.3923/std.2015.105.108.
  • Gigauri Kh., Abdaladze O., Nakhutsrishvili G., Akhalkatsi M. 2014. Vascular plant diversity and climate change in the alpine zone of the Central Greater Caucasus. International Journal of Ecosystems and Ecology Sciences (IJEES) 4 (4): 573-578.
  • Akhalkatsi M., Arabuli G., Lorenz R. Orchids as indicator species of forest disturbances on limestone quarry in Georgia (South Caucasus). J.Europäischer Orchideen 46 (1): 123-160.
  • Erschbammer B., Mallaun M., Unterluggauer P., Nakhutsrishvili G.Abdaladze O., Akhalkatsi M. Plant diversity along altitudinal gradients in the Central Alps and in the Central Greater Caucasus: The project GLORIA in south Tyrol (Italy) and in the Kasbegi region (Georgia). Ed. Pedrotti F., Stelvio ’70. Biodiversità, dinamica del paesaggio e gestione delle aree montane. Colloques phytosociologiques. Ediz. multilingue XXIX : 167-188.
  • Nakhutsrishvili G., Akhalkatsi M., Abdaladze O. First signs of transformation of alpine vegetation In the Caucasus as a result of climate warming (on the example of Kazbegi region). Ed. Pedrotti F., Stelvio ’70. Biodiversità, dinamica del paesaggio e gestione delle aree montane. Colloques phytosociologiques. Ediz. multilingue XXIX:405-414.
  • Lorenz R., Akhalkatsi M., Baumann H., Cortis P., Cogoni A., Scrugli A. Platanthera kuenkelei s.l. auf Sardinien und in Georgien, eine für Europa neue Artein Beitrag zu ihrer Taxonomie. Journal Europäischer Orchideen 44 (1): 3-62.
  • Otte A., Akhaltkatsi M., Nakhutsrishvili G., Simmering D., Waldhardt R. Phytodiversität in Georgien: Die Bedeutung von Standort und Landnutzung im Großen und Kleinen Kaukasus. Spiegel der Forschung 28 (2): 24-31.
  • Asanidze Z., Akhalkatsi M. Morphological Relationships between Wild and Cultivated Pears in Georgia. Proceedings of the Georgian Academy of Sciences Ser. Biology,  9 (1-4): 30-37.
  • Pistrick K., Akhalkatsi M., Girgvliani T., Shanshiashvili T. Sammlung von Saat- und Pflanzgut pflanzengenetischer Ressourcen in Svanetien und Lečchumi (Georgien, Kaukasus) 2009. Berichte Ges. Pflanzenbauwiss. 5 (5): 125-127.
  • Togonidze N., Akhalkatsi M. 2010. Sky exposition effect on phenological rhythm of Primula Proc. Tbilisi Bot. Gard. 98: 55-71 (Georg.).
  • Nakhutsrishvili G., Akhalkatsi M., Abdaladze O. Main Threats to the Mountain Biodiversity in Georgia (the Caucasus). Mountain Forum Bulletin 9 (2): 18-19.
  • Akhalkatsi M., Lorenz R., Mosulishvili M. Beiträge zur Kenntnis der Gattung Dactylorhiza in Georgien (Kaukasus). Ein Beitrag zum OPTIMA-Projekt »Kartierung der mediterranen Orchideen«.- Jber.  naturwiss. Ver. Wuppertal 60: 13-100.
  • Akhalkatsi M., Bokeria M., Maisaia I. Wild and Cultivated Species of Savory (Satureja L.) in Georgia and their Use. Proc. Tbilisi Botanical Garden 97: 126-131.
  • Maisaia I., Arabuli G., Akhalkatsi M., Mosulishvili M. Use of barley in being and folk medicine in mountain regions of Eastern and Southern Georgia. Proc. Tbilisi Bot. Gard. 96: 118-120 (Georg.).
  • Akhalkatsi M., Lorenz R., Mosulishvili M. 2006. Orchids and their habitats in Georgia. Journal Europäischer Orchideen 38 (2): 286 – 287.
  • Kimeridze M., Akhalkatsi M., Herbst P. Gefährdung und Schutzmaßnahmen für Georgiens Wälder. Forstzeitung 117 (4): 32-33.
  • Ekhvaia Zh., Akhalkatsi M. 2006. Comparative study of the quantitative parameters of berries and seeds in the autochthonous red grape varieties of the Kolkhis (Western Georgia). Proceedings of the Georgian Academy of Sciences Ser. Biology B, 4 (3): 38-46.
  • Nadirashvili N., Gvaladze G., Akhalkatsi M. 2006. Structure and function of the hypertrophic synergid in some species of genus Allium Proceedings of the Georgian Academy of Sciences Ser. Biology B, 4 (2): 53-60.
  • Akhalkatsi M., Baumann H., Lorenz R., Mosulishvili M., Peter R. Beiträge zur Kenntnis kaukasischer Orchideen. Journal Europäischer Orchideen 37 (4): 889-914.
  • Nadirashvili N., Gvaladze G., Akhalkatsi M. 2005. Atypical embryo sacs in Allium cepa Bulletin of the Georgian National Academy of Sciences, 171, 2:312-314.
  • Akhalkatsi M., Lorenz R., Matchutadze I., Mosulishvili M. Spiranthes amoena  a new species for flora of Georgia. Journal Europäischer Orchideen 36 (3): 745-754.
  • Akhalkatsi M., Gvaladze G., Gachechiladze M., Taralashvili N. Embryology of Gentiana angulosa and G. pontica (Gentianaceae). Proceedings of the Georgian Academy of Sciences Ser. Biology B 2 (1-2): 29-34.
  • Akhalkatsi M., Gvaladze G., Gachechiladze M. Relationships between structure of the reproductive organs and the life cycle in the family Gentianaceae. Bulletin of the Georgian National Academy of Sciences 170 (1): 125-129.
  • Nakhutsrishvili G., Abdaladze O., Akhalkatsi M. 2004. On the tree line vegetation in the Kazbegi region (the Central Caucasus). Bulletin of the Georgian National Academy of Sciences 169 (1): 124-129.
  • Nakhutsrishvili G., Abdaladze O., Akhalkatsi M. Global warming and treeline. Proceedings of the Georgian Academy of Sciences Ser. Biology B, 2 (1-2): 101-103.
  • Pistrick, M. Akhalkatsi, and G. Nakhutsrishvili. 2003. Collecting Allium in Georgia (Caucasus mountains) 2002. Allium Improvement Newsletter, 13: 1-7.
  • Gvaladze G., Nadirashvili N., Akhalkatsi M. 2002. Chromatin diminution during endosperm development in Allium atroviolaceum (Alliaceae). Bulletin of the Georgian National Academy of Sciences, 166 (3): 537-540.
  • Akhalkatsi M., Kimeridze M., Künkele S., Lorenz R., Mosulishvili M. Orchids of Georgia. Journal Europäischer Orchideen 33 (1): 457-458.
  • Akhalkatsi M., Gvaladze G. The ultrastructure of ovule sterile tissues of Peperomia  caperata (Piperaceae). Bulletin of the Georgian National Academy of Sciences 159 (1): 130-132.
  • Gvaladze G., Akhalkatsi M. 1998. Double fertilization in Peperomia pellucida. Phytomorphology 48 (4): 405-409.
  • Akhalkatsi M., Gvaladze G., Taralashvili N. Embryology of Primula algida and Primula amoena (Primulaceae). Bulletin of the Georgian National Academy of Sciences 157 (1): 98-100.
  • Gvaladze G., Akhalkatsi M. 1996. Comparative embryology of some Astragalus Bulletin of the Georgian National Academy of Sciences 153 (3): 432-434.
  • Wagner J., Achalkazi M., Mayr 1995. Anwendung quantitativ-embryologischer Methoden in Entwicklungsbiologie und Reproduktionsoekologie der Pflanzen. Anzeiger Oester. Akad. Wiss. math.-nat. Kl., 131: 7-18.
  • Gvaladze G., Chochia K., Akhalkatsi M. 1994. Fertilization in Indian lotus – Nelumbo nucifera (Nelumbonaceae). Bulletin of the Georgian National Academy of Sciences 150 (3): 515-518. (Russ.).
  • Vassilyev A.E., Akhalkatsi M. 1993. Structural and functional characterization of capitate trichoms in some Urticaceae. Botanicheskii Zhurnal (St. Petersburg) 78 (12): 1-8. (Russ.).
  • Akhalkatsi M., Gvaladze G. Seasonal changes of ultrastructure of embryo sac of Galanthus nivalis L. (Amaryllidaceae)Botanicheskii Zhurnal (St. Petersburg)  77 (1): 66-73. (Russ.).
  • Gvaladze G., Akhalkatsi M. On depression of chalazal part of an embryo sac in Angiosperms. Botanicheskii Zhurnal (St. Petersburg)  77 (7): 71-75. (Russ.).
  • Gvaladze G., Akhalkatsi M. 1990. Is the Polygonum type embryo sac primitive? Phytomorphology 40 (3-4): 331-337.
  • Akhalkatsi M., Beridze M., Gvaladze G. Female and male gametophytes of Astragalus caucasicus Pall. Proceedings of the Georgian Academy of Sciences Ser. Biology B 15 (4): 164-168. (Russ.).
  • Ciamporova M., Akhalkatsi M., Gvaladze G. Ultrastructure of the sterile tissues of the ovule of Galanthus nivalis L. (Amaryllidaceae). Botanicheskii Zhurnal (St. Petersburg) 73 (12): 1722-1730. (Russ.).
  • Gvaladze G., Akhalkatsi M. 1988. Ultrastructure of autumn and spring embryo sac of Galanthus nivalis Ann. Univ. L’ARERS, 23: 152-154.
  • Akhalkatsi M., Beridze M., Gvaladze G. Embryo and endosperm development in Astragalus caucasicus Pall. Bulletin of the Georgian National Academy of Sciences 132 (3): 601-604. (Georg).
  • Akhalkatsi M., Gvaladze G. Ultrastructure of the egg cell and zygote of Galanthus nivalis L. (Amaryllidaceae). Bull. Geor. Bot. Soc. 8: 187-199. (Russ.).

Articles in newspapers

  • Akhalkatsi M, Kimeridze M. Diversity of Georgian flora relationship of the Earth’s climate. Clean Environment 3: 3-25. (Georg.).
  • Akhalkatsi M, Kimeridze M. The effect of climate change in semi-arid regions in the reproduction of rare plants and diversity. Clean Environment 2: 3-30 (Georg.).
  • Akhalkatsi M., Kimeridze M. Implementation of the classification system of forest habitats in accordance with the ‘Natura2000’ standards in the Georgian Legislation. In proceeding of: Legal Aspects of European Forest Sustainable Development Proceedings of the 12th International Symposium Cyprus, At Lemesos, Cyprus, 11/2012; (Eds) Šulek, R., Herbst P. Schmithüsen F. Volume: IUFRO Division 9.
  • Ekhvaia J., Blattner F. R., Akhalkatsi M. 2010. Genetic diversity and relationships between wild grapevine (Vitis vinifera sylvestris) populations and aboriginal cultivars in Georgia. Proc. 33rd World Congress of Vine and Wine, 8th General Assambley of the OIV, 20-25 June 2010, Tbilisi, Georgia.
  • Akhalkatsi M., Ekhvaia J., Mosulishvili M., Nakhutsrishvili G., Abdaladze O., Batsatsashvili K. Erosion of crop genetic diversity in mountainous regions of Georgia. Mountain Diversity and Global Change. Spehn EM, Rudmann-Maurer K, Körner C, Maselli D (Eds): Conference of Parties of the CBD (COP10) in Nagoya, Japan in October 2010. GMBA-DIVERSITAS 2010.
  • Akhalkatsi M., Mosulishvili M., Kimeridze M., Maisaia I. Traditional use of medicinal plants in historical Tao-Klarjeti. Elkana, 1 (2): 4-6. (Georg.).
  • Akhalkatsi M., Fritsch R., Maisaia I., Nakhutsrishvili G., Pistrick K. Habitats of Allium Species in Georgia. In: Keusgen, M., Fritsch R.M. (eds.) “Botany, taxonomy and phytochemistry of wild Allium L. species of the Caucasus and Central Asia“. Marburg & Gatersleben, Germany,. pp. 45-52.
  • Pistrick K., Akhalkatsi M., Nakhutsrishvili G. Collecting Wild Allium Species in Georgia. In: Keusgen, M., Fritsch, R.M. (eds.), “Botany, taxonomy and phytochemistry of wild Allium L. species of the Caucasus and Central Asia“. Marburg & Gatersleben, Germany, pp. 147-156.
  • Kimeridze M., Akhalkatsi M., Herbst P. Gefährdung und Schutzmaßnahmen für Georgiens Wälder. In: Herbst, P., Mekić, F., Avdibegović, M., Schmithuesen, F. (eds). “Forstwirschaft und Forstrecht in den Reformstaaten Mittel- und Osteuropas 1990-2007”. Forstery Faculty Univ. Sarajevo, Bosnia/Herzegovina, pp. 31-34.
  • Kimeridze M., Akhalkatsi M. 2006. Forest Legislation in Relation with Biodiversity Conservation in Georgia. In: Schmithüsen, F., Herbst, P., Nonic, D., Jovic D., Stanisic M. (eds) “Legal Aspects of European Forest Sustainable Development”, pp.176-180, ETH,
  • Akhalkatsi M., Kimeridze M. Threats and Protection Measures of Forests in Georgia. In: Sedat, A., Yusuf, G., Aynur, A. C., Herbst P. (eds.) “Legal Aspects of European Forest Sustainable Development”. pp. 141-149, Department of Forest Law, Faculty of Forestry, Istanbul University.
  • Akhalkatsi M., Mosulishvili M., Kimeridze M., Maisaia I. Saffron- a valuable medicinal plant. Biomeurne, 1(12):33-35 (Georg.).
  • Akhalkatsi M., Kimeridze M., Maisaia I., Mosulishvili M. Flowless profits. CENN Caucasus Environmental NGO 4(13): 34-37.
  • Akhalkatsi M., Mosulishvili M., Kimeridze M. Iris family in Georgia. CENN Caucasus Environmental NGO 2 (7): 1316.
  • Akhalkatsi M., Mosulishvili M. Wetland orchid Spiranthes amoena in Georgia. CENN Caucasus Environmental NGO 1 (6): 30-31.
  • Akhalkatsi M., Kimeridze M., Kunkele S, Lorenz R., Mosulishvili M. Wild Orchids of Georgia. CENN Caucasus Environmental NGO 2 (3): 48-54.
  • Akhalkatsi M. 1998. Age and spatial structure in populations of three Astragalus species (Tragacantha, Fabaceae) In: Plant Life in High-Mountains. Eds. G. Nakhutsrishvili, O. Abdaladze, Tbilisi, pp. 20-26.

An extensive list of publications 

Current Courses

Course Catalog

  • BSc “Botany” (autumn), “Flawless profits: sustainable use of economic plants” (spring).
  • Practical BSc “Habitat identification” (spring), “Useful plants and fruits” (autumn and spring).
  • MSc “Ecosystems ecology” (autumn), “Wildlife and habitat biodiversity” (autumn), “Non-timber Forest and Nature Products, Management, Processing and Utilization” (spring).
  • MSc “Plant taxonomic features” (2006-2018);, “Plant reproductive ecology” (2006-2018);.
  • PhD course “Plant morphology and genetic resources”, Institute of Ecology, Ilia State University, Tbilisi (2010-2018).

  • 2012 ”Who’s Who in the World”. 29th ed. Marques Who’s Who publication board;
  • 2012 The Sir Isaac Newton Scientific Award of Excellence.
  • 2012 HeidelbergCement the quarry life award.
  • 2018   Citation: 1325;  h-index: 12; impact factor: 071.