Mumladze Levan
Ilia State University Room G 303, Giorgi Cereteli str. 3a, Tbilisi 0162, Georgia
Direction:Endocrinology and Pediatric Endocrinology
Position:Associate Professor
Dr. Mumladze received a PhD in Ecology at Ilia State University (2013). Since the establishment of Ilia State University, Dr. Mumladze has worked as an assistant dean at the Faculty of Life Science until 2010, as a research associate until 2013, as an Assistant professor until 2018, and as an Associate Professor associate from 2018 until now. Since 2003, Dr. Mumladze has been working in the Institute of Zoology in various positions, and since 2019, he has been serving as director of the Institute of Zoology. From 2022 he is an Editor-in-chief of the biodiversity research journal “Caucasiana (https://caucasiana.pensoft.net/).
Scientific interests / research interests
The scientific interests and current projects of Dr. Mumladze are related to biological diversity, distribution, and community composition. Based on model organisms (including molluscs, mites, several taxa of arthropods, and fishes), he is working to identify biodiversity patterns and structural mechanisms, interspecific relationships, biogeography and phylogeography in the Caucasus region.
Featured publications
- Murvanidze, M., Todria, N., Maraun, M., Mumladze, L., 2023. Annotated checklist of Georgian oribatid mites – II. Zootaxa 5227, 50–62. https://doi.org/https://doi.org/10.11646/zootaxa.5227.1.2
- Vilizzi, L., Piria, M., …… Mumladze, …… Zięba, G. 2022. Development and application of a multilingual electronic decision-support tool for risk screening non-native terrestrial animals under current and future climate conditions. NeoBiota 76, 211–236. https://doi.org/10.3897/neobiota.76.84268
- Pešić, V., Esen, Y., Gerecke, R., Goldschmidt, T., Mumladze, L., Smit, H., Zawal, A., 2022. Evidence of cryptic speciation in the Hygrobates calliger complex (Acariformes, Hydrachnidia, Hygrobatidae) with the description of two new species. Ecol. Montenegrina 59, 101–122. https://doi.org/https://dx.doi.org/10.37828/em.2022.59.10
- Mumladze, L., Kuljanishvili, T., Japoshvili, B., Epitashvili, G., Kalous, L., Vilizzi, L., Piria, M., 2022. Risk of invasiveness of non-native fishes in the South Caucasus biodiversity and geopolitical hotspot. NeoBiota 76, 109–133. https://doi.org/10.3897/neobiota.76.82776
- Maasri, A., Jähnig, S.C., …… Mumladze, …… Worischka, S. 2022. A global agenda for advancing freshwater biodiversity research. Ecol. Lett. 1–9. https://doi.org/10.1111/ele.13931
- Neiber, M.T., Walther, F., Kijashko, P. V, Mumladze, L., Hausdorf, B., 2022. The role of Anatolia in the origin of the Caucasus biodiversity hotspot illustrated by land snails in the genus Oxychilus. Cladistics 38, 83–102. https://doi.org/10.1111/cla.12479
- Bikashvili, A., Kachlishvili, N., Japoshvili, B., Mumladze, L., 2022. Species diversity and DNA barcode library of freshwater Molluscs of South Caucasus. Biodivers. Data J. 10, 1–24. https://doi.org/10.3897/BDJ.10.e84887
- Zerbe, S., Miller, R.M., Asanidze, Z., Asanidze, I., Mumladze, L., 2022. Local forest-related knowledge, perceptions, and perspectives as a basis for woodland restoration. For. Ecol. Landsc. Res. Nat. Conserv. 21, 31–43.
- Bakhtadze, N., Chakvetadze, N., Mumladze, L., Gabroshvili, N., 2021. Check-list of chromosome numbers of the family Hygromiidae (Gastropoda: Stylommatophora) with new data on Circassina frutis. Trav. du Muséum Natl. d’Histoire Nat. “Grigore Antipa” 64, 7–20. https://doi.org/10.3897/travaux.64.e69707
- Kuljanishvili, T., Mumladze, L., Japoshvili, B., Mustafayev, N., Ibrahimov, S., Patoka, J., Pipoyan, S., Kalous, L., 2021. The first unified inventory of non-native fishes of the South Caucasian countries, Armenia, Azerbaijan, and Georgia. Knowl. Manag. Aquat. Ecosyst. 422, 1–16. https://doi.org/10.1051/kmae/2021028
- Todria, N., Murvanidze, M., Mumladze, L., 2021. Oribatid (Acari: Oribatida) diversity in natural and altered open arid ecosystems of South-Eastern Caucasus. Pedobiologia (Jena). 87–88, 150750. https://doi.org/10.1016/j.pedobi.2021.150750
- Vilizzi, L., Copp, …… Mumladze, L., …… Clarke, S. 2021. A global-scale screening of non-native aquatic organisms to identify potentially invasive species under current and future climate conditions. Sci. Total Environ. 788, 147868. https://doi.org/10.1016/j.scitotenv.2021.147868
- Bikashvili, A., Kachlishvili, N., Mumladze, L., 2021. Species diversity and distribution of freshwater molluscs of Javakheti Highlands (Republic of Georgia). Biodivers. Data J. 9, e66649. https://doi.org/10.3897/BDJ.9.e66649
- Japoshvili, B., Couto, T.B.A., Mumladze, L., Epitashvili, G., McClain, M.E., Jenkins, C.N., Anderson, E.P., 2021. Hydropower development in the Republic of Georgia and implications for freshwater biodiversity conservation. Biol. Conserv. 263, 109359. https://doi.org/10.1016/j.biocon.2021.109359
- Neiber, M.T., Bikashvili, A., Bananashvili, G., Shubashishvili, A., Japoshvili, B., Walther, F., Mumladze, L., 2021. Continental molluscs collected during the second Georgian-German BioBlitz 2019 in Stepantsminda, Georgia. Mitteilungen der Dtsch. Malakozool. Gesellschaft 104, 23–36.
- Kuljanishvili, T., Japoshvili, B., Mumladze, L., Mustafayev, N., Ibrahimov, S., Patoka, J., Pipoyan, S., Kalous, L., 2021. Freshwater fish species diversity in Georgia (South Caucasus Region) and their local names, in: Kubík, Š., Barták, M. (Eds.), Proceedings of the „12th Workshop on Biodiversity”, Jevany. Jevany, Czech republic, pp. 48–65.
- Copp, G.H., Vilizzi, L., …… Mumladze, L., …… Mendoza, R. 2021. Speaking their language – Development of a multilingual decision-support tool for communicating invasive species risks to decision makers and stakeholders. Environ. Model. Softw. 135, 104900. https://doi.org/10.1016/j.envsoft.2020.104900
- Bláha, M., Patoka, J., Japoshvili, B., Let, M., Buřič, M., Kouba, A., Mumladze, L., 2021. Genetic diversity, phylogenetic position and morphometric analysis of Astacus colchicus (Decapoda, Astacidae): a new insight into Eastern European crayfish fauna. Integr. Zool. 16, 368–378. https://doi.org/10.1111/1749-4877.12493
- Kuljanishvili, T., Epitashvili, G., Freyhof, J., Japoshvili, B., Kalous, L., Levin, B., Mustafayev, N., Ibrahimov, S., Pipoyan, S., Mumladze, L., 2020. Checklist of the freshwater fishes of Armenia, Azerbaijan and Georgia. J. Appl. Ichthyol. 36, 501–514. https://doi.org/10.1111/jai.14038
- Epitashvili, G., Geiger, M.F., Astrin, J.J., Herder, F., Japoshvili, B., Mumladze, L., 2020. Towards retrieving the Promethean treasure: a first molecular assessment of the freshwater fish diversity of Georgia. Biodivers. Data J. e57862. https://doi.org/https://doi.org/10.3897/BDJ.8.e57862
- Copilaș-Ciocianu, D., Berci, G.M., Mumladze, L., 2020. First survey of shallow-water Amphipoda along the Georgian Black Sea coast reveals new faunistic records and the unexpected Atlantic invader Melita nitida. Mediterr. Mar. Sci. 21, 460–463. https://doi.org/10.12681/mms.22844
- Mumladze, L., Szekeres, M., 2020. A second extant species of Pontophaedusa Lindholm, 1924 (Gastropoda, Pulmonata, Clausiliidae) from Georgia. Ruthenica 30, 149–154.
- Pankvelashvili, E., Japoshvili, B., Mumladze, L., 2020. New remarkable record of Helicopsyche bacescui Orghidan and Botosaneanu, 1953 (Trichoptera, Helicopsychidae) from Georgia. Aquat. Insects 41, 184–188. https://doi.org/10.1080/01650424.2020.1737135
- Grego, J., Mumladze, L., Falniowski, A., Osikowski, A., Rysiewska, A., Palatov, D.M., Hofman, S., 2020. Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one n. Zookeys 955, 1–77. https://doi.org/10.3897/zookeys.955.51983
- Japoshvili, B., Lipinskaya, T., Gajduchenko, H., Sinchuk, A., Bikashvili, A., Mumladze, L., 2020. First DNA-based Records of New Alien Freshwater Species in the Republic of Georgia. Acta Zool. Bulg. 72, 545–551.
- Zerbe, S., Pieretti, L., Elsen, S., Asanidze, Z., Asanidze, I., Mumladze, L., 2020. Forest restoration potential in a deforested mountain area: an ecosociological approach towards sustainability. For. Sci. 66, 326–336. https://doi.org/10.1093/forsci/fxz081
- Mumladze, L., Japoshvili, B., Anderson, E.P., 2020. Faunal biodiversity research in the Republic of Georgia: a short review of trends, gaps, and needs in the Caucasus biodiversity hotspot. Biologia (Bratisl). 75, 1385–1397. https://doi.org/10.2478/s11756-019-00398-6
- Vrabec Vladimír , Berishvili Tamari , Kuljanishvili Tatia, Kulma Martin , Kulmová Karolína, B.T.& M.L., 2019. Observations on the composition of butterfly fauna in regions of Svaneti and Imereti, Georgia, in: Kubík, Š., Barták, M. (Eds.), 11th Workshop on Biodiversity. Jevany, Czech republic, pp. 175–189.
- Thormann, J., Ahrens, D., Anderson, C., Astrin, J.J., Mumladze, L., Rulik, B., Tarkhnishvili, D., Espeland, M., Geiger, M., Hein, N., Iankoshvili, G., Karalashvili, E., Mengual, X., Morkel, C., Neiber, M.T., Peters, R.S., Reimann, A., Ssymank, A., Wesener, T., Ziegler, J., Misof, B., 2019. A prelude to the Caucasus Barcode of Life Platform (CaBOL): Biodiversity Days in Georgia in 2018 and 2019. Bonn Zool. Bull. 68, 275–296. https://doi.org/10.20363/BZB-2019.68.2.275
- Nassirkhani, M., Mumladze, L., 2019. Redescription of Roncus crassipalpus Rafalski, 1949 (Pseudoscorpiones: Neobisiidae) from western Georgia. Arachnology 18, 133. https://doi.org/10.13156/arac.2018.18.2.133
- Nassirkhani, M., Zaragoza, J.A., Mumladze, L., 2019. A new pseudoscorpion genus from western Georgia (Pseudoscorpiones: Neobisiidae: Cornuroncus n. gen.), with a key to all Neobisiinae genera. Zootaxa 4624, 289–295. https://doi.org/http://dx.doi.org/10.11646/zootaxa.4624.2.12
- Maraun, M., Caruso, T., Hense, J., Lehmitz, R., Mumladze, L., Murvanidze, M., Nae, J., Schulz, J., Seniczak, A., Scheu, S., 2019. Parthenogenetic vs . sexual reproduction in oribatid mite communities. Ecol. Evol. 9, 7324–7332. https://doi.org/10.1002/ece3.5303
- Mumladze, L., Bikashvili, A., Kachlishvili, N., Grego, J., Japoshvili, B., Schniebs, K., Vinarski, M., Falniowski, A., Palatov, D., 2019. Progress towards research and conservation of Georgian freshwater molluscs. TENTACLE 27, 7–10.
- Murvanidze, M., Mumladze, L., Todria, N., Salakaia, M., 2019. Effect of ploughing and pesticide application on oribatid mite communities. Int. J. Acarol. 45, 181–188. https://doi.org/10.1080/01647954.2019.1572222
- Levin, B.A., Gandlin, A.A., Simonov, E.S., Levina, M.A., Barmintseva, A.E., Japoshvili, B., Mugue, N.S., Mumladze, L., Mustafayev, N.J., Pashkov, A.N., Roubenyan, H.R., Shapovalov, M., Doadrio, I., 2019. Phylogeny, phylogeography and hybridization of Caucasian barbels of the genus Barbus (Actinopterygii, Cyprinidae). Mol. Phylogenet. Evol. 135, 31–44. https://doi.org/10.1101/473173
- Nassirkhani, M., Mumladze, L., 2019. Neobisium (Neobisium) moreoticum (Pseudoscorpiones: Neobisiidae) from Georgia. Arachnol. Mitteilungen 57, 37–42. https://doi.org/10.30963/aramit5707
- Grego, J., Mumladze, L., 2019. Notes on vulnerability of the microendemic clausiliid species Acrotoma enguriensis from the southwestern Caucasus. TENTACLE 27, 27–29.
- Mumladze, L., Bikashvili, A., Japoshvili, B., Anistratenko, V. V, 2019. New alien species Mytilopsis leucophaeata and Corbicula fluminalis (Mollusca, Bivalvia) recorded in Georgia and notes on other non-indigenous molluscs invaded the South Caucasus. Vestn. Zool. 53, 187–194. https://doi.org/https://doi.org/10.2478/vzoo-2019-0019
- Japoshvili, G., Mumladze, L., 2019. New species and new records of Aphelinus Dalman (Hymenoptera: Chalcidoidea: Aphelinidae) from Lagodekhi Reserve (Sakartvelo-Georgia), with diversity and distribution along an elevational gradient. Turkish J. Zool. 43, 192–202. https://doi.org/10.3906/zoo-1802-3
- Shapovalov, M.I., Saprykin, M.A., Japoshvili, B., Mumladze, L., 2019. Materials to the fauna of aquatic bugs of the infraorder Nepomorpha (Heteroptera) of the Javakheti Highland, Georgia. Russ. Entomol. J. 28, 120–124. https://doi.org/10.15298/rusentj.28.2.02
- Nassirkhani, M., Mumladze, L., 2019. Notes on Roncus Microphthalmus (Daday, 1889) (Pseudoscorpiones: Neobisiidae), a polumorphic species, with a key to the Roncus species from the Middle East and the Caucasus region. Rev. Ibérica Aracnol. 34, 97–101.
- Todria, N., Murvanidze, M., Mumladze, L., 2019. Oribatid mite communities on former clay quarries under different reclamation strategy. Ann. Agrar. Sci. 17, 304–311.
- Murvanidze, M., Mumladze, L., Todria, N., 2019. A contribution to the knowledge of oribatid and mesostigmatic mites (Acari) with new records in Georgia. Persian J. Acarol. 8, 309–325. https://doi.org/http://dx.doi.org/10.22073/pja.v8i4.51419
- Murvanidze, M., Todria, N., Mumladze, L., & Kalatozishvili, L. (2018). Diversity of soil mite communities in different habitats of Saskhori quarries, Georgia. Persian Journal of Acarology, 7(3), 297–305. https://doi.org/10.22073/pja.v7i3.37647
- Nassirkhani, M., & Mumladze, L. (2018). A new cave pseudoscorpion (Pseudoscorpiones: Neobisiidae) from western Georgia. Arachnology, 17(9), 496-500. https://doi.org/10.13156/arac.2018.17.9.496
- Kuljanishvili, T., Mumladze, L., Kalous, L., & Japoshvili, B. (2018). Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia). Biologia, 73(1), 93-100. https://doi.org/10.2478/s11756-018-0012-y
- Gabelashvili, S., Mumladze, L., Bikashvili, A., Sroka, P., Godunko, R. J., & Japoshvili, B. (2018). The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country. Turkish Journal of Zoology, 42(2), 252-262. https://doi.org/10.3906/zoo-1709-4
- Mumladze L, Ulrich W, Asanidze Z, Japoshvili G (2017) An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea). Écoscience 24, 75–79. https://doi.org/10.1080/11956860.2017.1324717
- Mumladze L, Asanidze Z, Walther F, Hausdorf B (2017) Beyond elevation: Testing the climatic variability hypothesis vs. Rapoport’s rule in vascular plant and snail species in the Caucasus. Biol J Linn Soc 121, 753–763. https://doi.org/10.1093/biolinnean/blx027
- Grego J, Hofman S, Mumladze L, Falniowski A (2017) Agrafia Szarowska et Falniowski, 2011 (Caenogastropoda: Hydrobiidae) in the Caucasus. Folia Malacologica 25, 237–247. https://doi.org/10.12657/folmal.025.025
- Mumladze L, Murvanidze M, Maraun M (2017). Patterns of oribatid mite species diversity: testing the effects of elevation, area and sampling effort. Experimental and applied acarology 72(3), 245-262. https://doi.org/10.1007/s10493-017-0153-7
- Aslan EG, Mumladze L, Japoshvili G (2017) List of leaf beetles (Coleoptera: Chrysomelidae) from Lagodekhi reserve with new records for Transcaucasia and Georgia. Zootaxa 4277, 86-98. https://doi.org/10.11646/zootaxa.4277.1.6
- Japoshvili B, Mumladze L, Murvanidze L (2017) The population of Carassius gibelio (Bloch, 1782) and its parasites in Madatapa Lake (South Georgia). Iranian Journal of Fisheries Sciences 16(2), 793-799.
- Bakhtadze NG, Chakvetadze NL, Mumladze LJ, Bakhtadze GI, Tskhadaia EA (2016) Karyological data of terrestrial mollusks (Mollusca: Gastropoda: Pulmonata) of Georgia. Proceedings of the Institute of Zoology XXV: 23-27.
- Gabelashvili S, Bikashvili A, Shubitidze Z, Gioshvili M, Pankvelashvili E, Mumladze L, Japoshvili B (2016) Family level diversity and distribution of macroinvertebrates of Madatapa, Khanchali and Bughdasheni lakes in Javakheti plateau (South Georgia). Proceedings of the Institute of Zoology XXV: 117-127.
- Mumladze L, Paposhvili N (2016) A New Addition to the Malacofauna of Georgia – Eobania Vermiculata is Replenishing its Range. Proceedings of the Institute of Zoology XXV:153-155.
- Seehausen M, Schröter A, Mumladze L, Grebe B (2016) Additional Odonata records from Georgia, southern Caucasus ecoregion,with the first record of Ischnura fountaineae (Odonata:Coenagrionidae). Notulae Odonatologicae 8 (8), 266-283. https://doi.org/
- Murvanidze M, Mumladze L, Arabuli T, Barjadze S, Salakaia M, (2016) Oribatida diversity in different microhabitats of Mtirala National Park. Journal of Acarological Society of Japan 25(S1), 35-49.
- Tarkhnishvili D, Gabelaia M, Mumladze L, Murtskhvaladze M, (2016) Mitochondrial phylogeny of the Darevskia saxicola complex: two highly deviant evolutionary lineages from the easternmost part of the range. Herpetological Journal 26, 175–182.
- Murvanidze M, Mumladze L (2016) Annotated checklist of Georgian oribatid mites. Zootaxa 4089 (1), 1-81. https://doi.org/10.11646/zootaxa.4089.1
- Mumladze L (2015) Species of the Genus Helix (Mollusca, Gastropoda) in Georgia. Proceedings of the Institute of Zoology XXIV, 148-157.
- Japoshvili G, Fallahzadeh M, Mumladze L (2015) Predictive modeling for better understanding distribution and systematics of useful Encyrtids (Hymenotpera: Chalcidoidea: Encyrtidae). Annals of Agrarian Science 13(1), 33-42.
- Barjadze S, Murvanidze M, Arabuli T, Mumladze L, Pkhakadze V, Djanashvili R, Salakaia M (2015) Annotated list of invertebrates of the Georgian karst caves. Georgian Academic Book, Tbilisi. ISBN: 978- 9941-9371-2-5.
- Mumladze L, Murvanidze M, Maraun M, Salakaia M (2015) Oribatid mite communities along an elevational gradient in Sairme gorge (Caucasus). Experimental and Applied Acarology 66(1), 41-51. https://doi.org/10.1007/s10493-015-9893-4
- Mumladze L (2014) Sympatry without co-occurrence: exploring the pattern of distribution of two Helix species in Georgia using an ecological niche modelling approach. Journal of Molluscan Studies 80(3), 249-255. https://doi.org/10.1093/mollus/eyu045
- Walther F, Kijashko PV, Harutyunova L, Mumladze L, Neiber MT, Hausdorf B (2014) Biogeography of the land snails of the Caucasus region. TENTACLE 3-5.
- Murvanidze M, Mumladze L (2014) Oribatid mite (Acari: Oribatida) diversity in different forest stands of Borjom-Kharagauli National Park (Georgia). Persian Journal of Acarology 3(4), 257–276. https://doi.org/
- Bakhtadze NG, Chakvetadze NL, Mumladze LJ, Bakhtadze GI, Tskhadaia EA (2014) Karyological Studies in Terrestrial Mollusks (Mollusca: Gastropoda: Pulmonata) of Georgia. International Conference «Biological Diversity and Conservation Problems of the Fauna of the Caucasus – 2», Armenia pp59-63.
- Mumladze L, Cameron RAD, Pokryszko B (2014) Endemic Land Molluscs in Georgia: How Well are they Protected by Existing Reserves and National Parks? Journal of Molluscan Studies 80, 67-73. https://doi.org/10.1093/mollus/eyt047
- Japoshvili B, Mumladze L, Kucuk F (2013) Invasive Carassius Carp in Georgia: Current State of Knowledge and Future Perspectives. Current Zoology 59, 732-739. https://doi.org/
- Murvanidze M, Mumladze M, Arabuli T, Kvavadze E (2013) Oribatid mite colonization of sandand manganese tailing sites. Acarologia 53, 203-215. https://doi.org/10.1051/acarologia/20132089
- Mumladze L, Tarkhnishvili D, Murtskhvaladze M (2013) Systematics and Evolutionary Hystori of Large Endemic Snails from the Caucasus (Helix buchii, H. goderdzianai (Helicidae). American Malacological Bulletin 31, 225-234. https://doi.org/10.4003/006.031.0202
- Mumladze L (2013) Shell size differences in Helix lucorum Linnaeus, 1758 (Mollusca: Gastropoda) between natural and urban environments. Turkish Journal of Zoology 37, 1-6. https://doi.org/10.3906/zoo-1206-10
- Mumladze L, Murvanidze M, Behan-Pelletier V (2013) Compositional patterns in Holarctic peat bog inhabiting oribatid mite (Acari: Oribatida) communities. Pedobiologia 56, 41-48. https://doi.org/10.1016/j.pedobi.2012.10.001
- Tarkhnishvili D, Gavashelishvili A, Mumladze L (2012) Palaeoclimatic models help to understand currentdistribution of Caucasian forest species. Biological Journal of the Linean Society 105, 231-248. https://doi.org/10.1111/j.1095-8312.2011.01788.x
- Pokryszko BM, Cameron RAD, Mumladze L, Tarkhnishvili D (2011) Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium. Biological Journal of the Linean Society 102, 239-250. https://doi.org/10.1111/j.1095-8312.2010.01575.x
- Murvanidze M, Kvavadze E, Mumladze L, Arabuli T (2011) Comparison of Earthworms (Lumbricidae) and Oribatid Mite (Acari, Oribatida) communities in Natural and Urban Ecosystems. Vestnik zoologii 45, 16-24. https://doi.org/10.2478/v10058-011-0021-6
- Murvanidze M, Mumladze L, Arabuli T, Kvavadze E (2011) Landscape distribution of oribatid mites (Acari,Oribatida) in Colchis National Park (Georgia, Caucasus). Zoosymposia 6, 221-233. https://doi.org/10.11646/zoosymposia.6.1.32
- Tarkhnishvili D, Gavashelishvili A, Avaliani A, Murtskhvaladze M, Mumladze L (2010) Unisexual rocklizard might be outcompeting its bisexual progenitors in the Caucasus. Biological Journal of the Linnean Society 101, 447–460. https://doi.org/10.1111/j.1095-8312.2010.01498.x
- Mumladze L, Tarkhnishvili D, Pokryszko BM (2008) A new species of the genus Helix from the Lesser Caucasus (SW Georgia). Journal of Conchology 39, 483-485
- Kvavadze EV, Rukhadze L, Nikolaishvili V, Mumladze L. (2008) Botanical and zoological remains from an earlymedieval grave at Tsitsamuri, Georgia. Vegetation History and Archaeobotany 17, 217-224. https://doi.org/10.1007/s00334-008-0183-5
- Murvanidze M, Arabuli T, Kvavadze ES, Mumladze L (2008) The Effect of Fire Disturbance on Oribatid Mite Communities. Integrative Acarology. Proceedings of the 6th European Congress. European Association of Acarolgists pp. 216-221
Current Courses |
Course Catalog |
|---|---|
| Master’s level
Bachelor’s level
Master’s level
|
2019-2020 Year Bachelor’s level
Master’s level
Doctoral level
2019-2020 Year Bachelor’s level
Master’s level
Doctoral level
2018-2019 Year Bachelor’s level
Doctoral level
2017-2018 Year Bachelor’s level
Master’s level
2016-2017 Year Bachelor’s level
Master’s level
2015-2016 Year Bachelor’s level
Master’s level
2014-2015 Year Master’s level
2013-2014 Year Master’s level
2010-2011 Year უმ.პროფესიული
2009-2010 Year Bachelor’s level
|
https://sites.google.com/view/mumladze/home?authuser=0